0000001 - Document - Cover link:presentationLink link:calculationLink link:definitionLink 0000002 - Statement - Condensed Consolidated Balance Sheets link:presentationLink link:calculationLink link:definitionLink 0000003 - Statement - Condensed Consolidated Balance Sheets (Parenthetical) link:presentationLink link:calculationLink link:definitionLink 0000004 - Statement - Condensed Consolidated Statements of Operations link:presentationLink link:calculationLink link:definitionLink 0000005 - Statement - Condensed Consolidated Statements of Comprehensive Loss link:presentationLink link:calculationLink link:definitionLink 0000006 - Statement - Condensed Consolidated Statements of Stockholders' Equity link:presentationLink link:calculationLink link:definitionLink 0000007 - Statement - Condensed Consolidated Statements of Cash Flows link:presentationLink link:calculationLink link:definitionLink 0000008 - Disclosure - Organization link:presentationLink link:calculationLink link:definitionLink 0000009 - Disclosure - Summary of Significant Accounting Policies link:presentationLink link:calculationLink link:definitionLink 0000010 - Disclosure - Collaboration Agreements link:presentationLink link:calculationLink link:definitionLink 0000011 - Disclosure - Condensed Consolidated Balance Sheet Components link:presentationLink link:calculationLink link:definitionLink 0000012 - Disclosure - Fair Value Measurements link:presentationLink link:calculationLink link:definitionLink 0000013 - Disclosure - Commitments and Contingencies link:presentationLink link:calculationLink link:definitionLink 0000014 - Disclosure - Common Stock link:presentationLink link:calculationLink link:definitionLink 0000015 - Disclosure - Stock-Based Compensation link:presentationLink link:calculationLink link:definitionLink 0000016 - Disclosure - Defined Contribution Plan link:presentationLink link:calculationLink link:definitionLink 0000017 - Disclosure - Income Taxes link:presentationLink link:calculationLink link:definitionLink 0000018 - Disclosure - Net Loss Per Share link:presentationLink link:calculationLink link:definitionLink 0000019 - Disclosure - Related Party Transactions link:presentationLink link:calculationLink link:definitionLink 0000020 - Disclosure - Subsequent Events link:presentationLink link:calculationLink link:definitionLink 0000021 - Disclosure - Summary of Significant Accounting Policies (Policies) link:presentationLink link:calculationLink link:definitionLink 0000022 - Disclosure - Condensed Consolidated Balance Sheet Components (Tables) link:presentationLink link:calculationLink link:definitionLink 0000023 - Disclosure - Fair Value Measurements (Tables) link:presentationLink link:calculationLink link:definitionLink 0000024 - Disclosure - Commitments and Contingencies (Tables) link:presentationLink link:calculationLink link:definitionLink 0000025 - Disclosure - Common Stock (Tables) link:presentationLink link:calculationLink link:definitionLink 0000026 - Disclosure - Stock-Based Compensation (Tables) link:presentationLink link:calculationLink link:definitionLink 0000027 - Disclosure - Net Loss Per Share (Tables) link:presentationLink link:calculationLink link:definitionLink 0000028 - Disclosure - Organization - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 0000029 - Disclosure - Collaboration Agreements - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 0000030 - Disclosure - Condensed Consolidated Balance Sheet Components - Summary of Property and Equipment (Details) link:presentationLink link:calculationLink link:definitionLink 0000031 - Disclosure - Condensed Consolidated Balance Sheet Components - Schedule of Accrued Expenses and Other Current Liabilities (Details) link:presentationLink link:calculationLink link:definitionLink 0000032 - Disclosure - Fair Value Measurements - Summary of Investments Fair Value Measurement on Recurring Basis (Details) link:presentationLink link:calculationLink link:definitionLink 0000033 - Disclosure - Fair Value Measurements - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 0000034 - Disclosure - Commitments and Contingencies - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 0000035 - Disclosure - Commitments and Contingencies - Other Information Related to Leases (Details) link:presentationLink link:calculationLink link:definitionLink 0000036 - Disclosure - Common Stock - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 0000037 - Disclosure - Common Stock - Schedule of Common Stock Shares Reserved for Future Issuance (Details) link:presentationLink link:calculationLink link:definitionLink 0000038 - Disclosure - Stock-Based Compensation - Schedule of Option Activity under the Stock Plans (Details) link:presentationLink link:calculationLink link:definitionLink 0000039 - Disclosure - Stock-Based Compensation - Schedule of RSU Activity (Details) link:presentationLink link:calculationLink link:definitionLink 0000040 - Disclosure - Stock-Based Compensation - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 0000041 - Disclosure - Stock-Based Compensation - Summary of Stock-Based Compensation Expense Related to Stock Options and ESPP Included in the Statements of Operations (Details) link:presentationLink link:calculationLink link:definitionLink 0000042 - Disclosure - Defined Contribution Plan - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 0000043 - Disclosure - Income Taxes - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 0000044 - Disclosure - Net Loss Per Share - Schedule of Computation of Basic and Diluted Net Loss per Share Attributable to Common Stockholders (Details) link:presentationLink link:calculationLink link:definitionLink 0000045 - Disclosure - Net Loss Per Share - Schedule of Antidilutive Securities Excluded from Computation of Diluted Net Loss per Share of Common Stock (Details) link:presentationLink link:calculationLink link:definitionLink 0000046 - Disclosure - Related Party Transactions - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink