0000001 - Document - Cover link:presentationLink link:calculationLink link:definitionLink 0000002 - Document - Audit Information link:presentationLink link:calculationLink link:definitionLink 0000003 - Statement - CONSOLIDATED BALANCE SHEETS link:presentationLink link:calculationLink link:definitionLink 0000004 - Statement - CONSOLIDATED BALANCE SHEETS (Parenthetical) link:presentationLink link:calculationLink link:definitionLink 0000005 - Statement - CONSOLIDATED STATEMENTS OF OPERATIONS link:presentationLink link:calculationLink link:definitionLink 0000006 - Statement - CONSOLIDATED STATEMENTS OF COMPREHENSIVE LOSS link:presentationLink link:calculationLink link:definitionLink 0000007 - Statement - CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY link:presentationLink link:calculationLink link:definitionLink 0000008 - Statement - CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY (Parenthetical) link:presentationLink link:calculationLink link:definitionLink 0000009 - Statement - CONSOLIDATED STATEMENTS OF CASH FLOWS link:presentationLink link:calculationLink link:definitionLink 0000010 - Disclosure - Organization link:presentationLink link:calculationLink link:definitionLink 0000011 - Disclosure - Summary of Significant Accounting Policies link:presentationLink link:calculationLink link:definitionLink 0000012 - Disclosure - Collaboration Agreements link:presentationLink link:calculationLink link:definitionLink 0000013 - Disclosure - Consolidated Balance Sheet Components link:presentationLink link:calculationLink link:definitionLink 0000014 - Disclosure - Fair Value Measurements link:presentationLink link:calculationLink link:definitionLink 0000015 - Disclosure - Commitments and Contingencies link:presentationLink link:calculationLink link:definitionLink 0000016 - Disclosure - Common Stock link:presentationLink link:calculationLink link:definitionLink 0000017 - Disclosure - Stock-Based Compensation link:presentationLink link:calculationLink link:definitionLink 0000018 - Disclosure - Defined Contribution Plan link:presentationLink link:calculationLink link:definitionLink 0000019 - Disclosure - Income Taxes link:presentationLink link:calculationLink link:definitionLink 0000020 - Disclosure - Net Loss Per Share link:presentationLink link:calculationLink link:definitionLink 0000021 - Disclosure - Related Party Transactions link:presentationLink link:calculationLink link:definitionLink 9954471 - Disclosure - Summary of Significant Accounting Policies (Policies) link:presentationLink link:calculationLink link:definitionLink 9954472 - Disclosure - Consolidated Balance Sheet Components (Tables) link:presentationLink link:calculationLink link:definitionLink 9954473 - Disclosure - Fair Value Measurements (Tables) link:presentationLink link:calculationLink link:definitionLink 9954474 - Disclosure - Commitments and Contingencies (Tables) link:presentationLink link:calculationLink link:definitionLink 9954475 - Disclosure - Common Stock (Tables) link:presentationLink link:calculationLink link:definitionLink 9954476 - Disclosure - Stock-Based Compensation (Tables) link:presentationLink link:calculationLink link:definitionLink 9954477 - 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Commitments and Contingencies - Schedule of Undiscounted Future Non-cancellable Lease Payments (Details) link:presentationLink link:calculationLink link:definitionLink 9954490 - Disclosure - Common Stock - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9954491 - Disclosure - Common Stock - Schedule of Common Stock Shares Reserved for Future Issuance (Details) link:presentationLink link:calculationLink link:definitionLink 9954492 - Disclosure - Stock-Based Compensation - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9954493 - Disclosure - Stock-Based Compensation - Schedule of Option Activity Under the Stock Plans (Details) link:presentationLink link:calculationLink link:definitionLink 9954494 - Disclosure - Stock-Based Compensation - Schedule of Option Activity Under the Stock Plans Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9954495 - Disclosure - Stock-Based Compensation - Summary of Estimated Fair Value of Stock Options and ESPP Using the Assumptions (Details) link:presentationLink link:calculationLink link:definitionLink 9954496 - Disclosure - Stock-Based Compensation - Schedule of RSU Activity (Details) link:presentationLink link:calculationLink link:definitionLink 9954497 - Disclosure - Stock-Based Compensation - Summary of Stock-Based Compensation Expense Related to Stock Options RSUs and ESPP Included in Statements of Operations (Details) link:presentationLink link:calculationLink link:definitionLink 9954498 - Disclosure - Defined Contribution Plan - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9954499 - Disclosure - Income Taxes - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9954500 - Disclosure - Income Taxes - Components of Loss Before Income Taxes (Details) link:presentationLink link:calculationLink link:definitionLink 9954501 - Disclosure - Income Taxes - Effective Tax Rate Differs from Federal Statutory Rate (Details) link:presentationLink link:calculationLink link:definitionLink 9954502 - Disclosure - Income Taxes - Significant Components of Deferred Tax Assets for Federal and State Income Taxes (Details) link:presentationLink link:calculationLink link:definitionLink 9954503 - Disclosure - Income Taxes - Reconciliation of Beginning and Ending Amounts of Unrecognized Income Tax Benefits (Details) link:presentationLink link:calculationLink link:definitionLink 9954504 - Disclosure - Net Loss Per Share - Schedule of Computation of Basic and Diluted Net Loss per Share (Details) link:presentationLink link:calculationLink link:definitionLink 9954505 - Disclosure - Net Loss Per Share - Schedule of Computation of Basic and Diluted Net Loss per Share Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9954506 - Disclosure - Net Loss Per Share - Schedule of Antidilutive Securities Excluded from Computation of Diluted Net Loss per Share of Common Stock (Details) link:presentationLink link:calculationLink link:definitionLink 9954507 - Disclosure - Related Party Transactions - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink