0000001 - Document - Cover link:presentationLink link:calculationLink link:definitionLink 0000002 - Document - Audit Information link:presentationLink link:calculationLink link:definitionLink 9952151 - Statement - CONSOLIDATED BALANCE SHEETS link:presentationLink link:calculationLink link:definitionLink 9952152 - Statement - CONSOLIDATED BALANCE SHEETS (Parenthetical) link:presentationLink link:calculationLink link:definitionLink 9952153 - Statement - CONSOLIDATED STATEMENTS OF OPERATIONS link:presentationLink link:calculationLink link:definitionLink 9952154 - Statement - CONSOLIDATED STATEMENTS OF COMPREHENSIVE LOSS link:presentationLink link:calculationLink link:definitionLink 9952155 - Statement - CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY link:presentationLink link:calculationLink link:definitionLink 9952156 - Statement - CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY (Parenthetical) link:presentationLink link:calculationLink link:definitionLink 9952157 - Statement - CONSOLIDATED STATEMENTS OF CASH FLOWS link:presentationLink link:calculationLink link:definitionLink 9952158 - Disclosure - Organization link:presentationLink link:calculationLink link:definitionLink 9952159 - Disclosure - Summary of Significant Accounting Policies link:presentationLink link:calculationLink link:definitionLink 9952160 - Disclosure - Collaboration Agreements link:presentationLink link:calculationLink link:definitionLink 9952161 - Disclosure - Consolidated Balance Sheet Components link:presentationLink link:calculationLink link:definitionLink 9952162 - Disclosure - Fair Value Measurements link:presentationLink link:calculationLink link:definitionLink 9952163 - Disclosure - Commitments and Contingencies link:presentationLink link:calculationLink link:definitionLink 9952164 - Disclosure - Common Stock link:presentationLink link:calculationLink link:definitionLink 9952165 - Disclosure - Stock-Based Compensation link:presentationLink link:calculationLink link:definitionLink 9952166 - Disclosure - Defined Contribution Plan link:presentationLink link:calculationLink link:definitionLink 9952167 - Disclosure - Income Taxes link:presentationLink link:calculationLink link:definitionLink 9952168 - Disclosure - Net Loss Per Share link:presentationLink link:calculationLink link:definitionLink 9952169 - Disclosure - Related Party Transactions link:presentationLink link:calculationLink link:definitionLink 9955511 - Disclosure - Summary of Significant Accounting Policies (Policies) link:presentationLink link:calculationLink link:definitionLink 9955512 - Disclosure - Consolidated Balance Sheet Components (Tables) link:presentationLink link:calculationLink link:definitionLink 9955513 - Disclosure - Fair Value Measurements (Tables) link:presentationLink link:calculationLink link:definitionLink 9955514 - Disclosure - Commitments and Contingencies (Tables) link:presentationLink link:calculationLink link:definitionLink 9955515 - Disclosure - Common Stock (Tables) link:presentationLink link:calculationLink link:definitionLink 9955516 - Disclosure - Stock-Based Compensation (Tables) link:presentationLink link:calculationLink link:definitionLink 9955517 - Disclosure - Income Taxes (Tables) link:presentationLink link:calculationLink link:definitionLink 9955518 - Disclosure - Net Loss Per Share (Tables) link:presentationLink link:calculationLink link:definitionLink 9955519 - Disclosure - Organization (Details) link:presentationLink link:calculationLink link:definitionLink 9955520 - Disclosure - Summary of Significant Accounting Policies (Details) link:presentationLink link:calculationLink link:definitionLink 9955521 - Disclosure - Collaboration Agreements (Details) link:presentationLink link:calculationLink link:definitionLink 9955522 - Disclosure - Consolidated Balance Sheet Components - Summary of Property and Equipment (Details) link:presentationLink link:calculationLink link:definitionLink 9955523 - Disclosure - Consolidated Balance Sheet Components - Summary of Accrued Expenses and Other Current Liabilities (Details) link:presentationLink link:calculationLink link:definitionLink 9955524 - Disclosure - Fair Value Measurements - Schedule of Investments Fair Value Measurement on Recurring Basis (Details) link:presentationLink link:calculationLink link:definitionLink 9955525 - Disclosure - Fair Value Measurements - AFS Unrealized Position (Details) link:presentationLink link:calculationLink link:definitionLink 9955526 - Disclosure - Fair Value Measurements - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9955527 - Disclosure - Commitments and Contingencies - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9955528 - Disclosure - Commitments and Contingencies - Schedule of Other Information Related to Leases (Details) link:presentationLink link:calculationLink link:definitionLink 9955529 - Disclosure - Commitments and Contingencies - Schedule of Undiscounted Future Non-cancellable Lease Payments (Details) link:presentationLink link:calculationLink link:definitionLink 9955529 - Disclosure - Commitments and Contingencies - Schedule of Undiscounted Future Non-cancellable Lease Payments (Details) link:presentationLink link:calculationLink link:definitionLink 9955530 - Disclosure - Common Stock - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9955531 - Disclosure - Common Stock - Schedule of Common Stock Shares Reserved for Future Issuance (Details) link:presentationLink link:calculationLink link:definitionLink 9955532 - Disclosure - Stock-Based Compensation - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9955533 - Disclosure - Stock-Based Compensation - Schedule of Option Activity Under the Stock Plans (Details) link:presentationLink link:calculationLink link:definitionLink 9955534 - Disclosure - Stock-Based Compensation - Summary of Estimated Fair Value of Stock Options and ESPP Using the Assumptions (Details) link:presentationLink link:calculationLink link:definitionLink 9955535 - Disclosure - Stock-Based Compensation - Schedule of RSU Activity (Details) link:presentationLink link:calculationLink link:definitionLink 9955536 - Disclosure - Stock-Based Compensation - Summary of Stock-Based Compensation Expense Related to Stock Options RSUs and ESPP Included in Statements of Operations (Details) link:presentationLink link:calculationLink link:definitionLink 9955537 - Disclosure - Defined Contribution Plan (Details) link:presentationLink link:calculationLink link:definitionLink 9955538 - Disclosure - Income Taxes - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9955539 - Disclosure - Income Taxes - Components of Loss Before Income Taxes (Details) link:presentationLink link:calculationLink link:definitionLink 9955540 - Disclosure - Income Taxes - Provision for Income Taxes (Details) link:presentationLink link:calculationLink link:definitionLink 9955541 - Disclosure - Income Taxes - Effective Tax Rate Differs from Federal Statutory Rate (Details) link:presentationLink link:calculationLink link:definitionLink 9955542 - Disclosure - Income Taxes - Significant Components of Deferred Tax Assets for Federal and State Income Taxes (Details) link:presentationLink link:calculationLink link:definitionLink 9955543 - Disclosure - Income Taxes - Reconciliation of Beginning and Ending Amounts of Unrecognized Income Tax Benefits (Details) link:presentationLink link:calculationLink link:definitionLink 9955544 - Disclosure - Net Loss Per Share - Schedule of Computation of Basic and Diluted Net Loss per Share (Details) link:presentationLink link:calculationLink link:definitionLink 9955545 - Disclosure - Net Loss Per Share - Schedule of Antidilutive Securities Excluded from Computation of Diluted Net Loss per Share of Common Stock (Details) link:presentationLink link:calculationLink link:definitionLink 9955546 - Disclosure - Related Party Transactions (Details) link:presentationLink link:calculationLink link:definitionLink